One would have to know all the more relevant physical and biotic factors and be able to depict their interactions . 人们必须知道所有有关的物理因子和生物因子，并能描述其相互影响。
2.
Showed a distinct seasonality . both abiotic and biotic factors were also found to influence the growth of the seagrass species 而生物性及非生物性的因素也被发现会影响海草的成长。
